Ph.D. Developmental, cellular, and Molecular Biology, University of Texas at Austin, Austin, Texas, U.S.A (1988)
Dissertation title: “Use of a Bioassay to Characterize Anterior Cytoplasmic Determinants in Chironomus Samoensis Eggs”
M.Sc. Biology, Loyola Marymount University, L.A. California, U.S.A (1981)
B.Sc. Zoology, Alexandria University, Alexandria, Egypt (1977).
